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Tool and editing shortcuts apply while a project is open and an input, textarea, or editable text region is not focused.

Tools

Change tool characters in Settings → Shortcuts. Releasing Space restores the previous tool. Switching away from Koharu cancels an unfinished canvas gesture.

Editing

Use a unique character for each tool. If bindings conflict, the first matching tool in toolbar order wins.
Shortcuts affect the desktop interface, not the exported page.