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Tool and editing shortcuts apply while a project is open and an input, textarea, or editable text region is not focused.

Tools

Change tool characters in Settings → Shortcuts. The active tool does not change while Space is held. Switching away from Koharu cancels an unfinished canvas gesture.

Editing

Use a unique character for each tool. If bindings conflict, the Fit Window key is checked first, then the first matching tool in toolbar order wins. Custom bindings reset when Koharu restarts.
Shortcuts affect the desktop interface, not the exported page.